The Anthonomus squamosus species-group (Coleoptera: Curculionidae: Curculioninae: Anthonomini) contains 31 species in North America. The known host plants of the group are divided between species of Asteraceae and Malvaceae. Characters diagnostic of the group and descriptions of each of the species are given and illustrated, and a key to the species is presented. Host plant associations are given, when known. Seven new species are described: Anthonomus proximatus Clark and Burke, new species (USA: AR, MI, OK, TN); Anthonomus vittatus Clark and Burke, new species (USA: CA, UT); Anthonomus smithi Clark and Burke, new species (Mexico: Chiapas, Colima, Guerrero, Hidalgo, Jalisco, Michoacán, Morelos, Nayarit, Oaxaca, Puebla, San Luis Potosí, Sonora, Veracruz); Anthonomus obrienorum Clark and Burke, new species (USA: CA); Anthonomus bowditchi Clark and Burke, new species, (Mexico: Sonora, Veracruz); Anthonomus robinsonelli Clark and Burke, new species (Mexico: Chiapas, Hidalgo, Nuevo León, Tamaulipas); and Anthonomus curvicrus Clark and Burke, new species (Mexico: Estado de México, Oaxaca, Tlaxcala, Veracruz). Lectotypes are designated for Anthonomus squamans Champion, Anthonomus solarii Champion, Anthonomus tectus LeConte, Anthonomus testaceosquamosus Linell, and Anthonomus rubricosus Boheman.

Five new species of the genus Pandeleteius Schönherr from South America are described: Pandeleteius emarginatus Girón and Howden, new species, from Brazil; Pandeleteius genieri Girón and Howden, new species, from Brazil and Peru; Pandeleteius obrienorum Girón and Howden, new species, from Ecuador and Peru; Pandeleteius pavo Girón and Howden, new species, from Peru; and Pandeleteius tupi Girón and Howden, new species, from Brazil. Photographs of the habitus and genitalia and a distribution map are presented along with a list of the species of Pandeleteius known from Brazil, Ecuador, and Peru. This paper constitutes the last contribution of Anne T. Howden on the taxonomy and biodiversity of the genus Pandeleteius.

The genus AporiusKuschel, 1955, described based on the single species Hadropus brevipennisChevrolat, 1879 from French Guiana, is herein transferred from the tribe Tanymecini, subtribe Piazomiina to the tribe Naupactini, new tribal placement, because it shares the typical characters of this tribe, except the presence of connate tarsal claws. Within Naupactini, Aporius is probably related to Hadropus Schoenherr, 1826, based on the morphology of the rostrum, antennae, and genitalia. We redescribe and illustrate the genus Aporius and its single species, and we provide a key to separate it from Hadropus. We also discuss the taxonomic status of the remaining members of South American Piazomiina and assign the genus Galenactus Kuschel to Leptopiini, new tribal placement, thus removing Piazomiina from occurrence in South America.

Kosmimodes howdenae Setliff, new genus and new species, is described for a brilliant metallic green broad-nosed weevil from the dry forests of Pedernales Province in the Dominican Republic. Kosmimodes is closely related to Apotomoderes Dejean and Melathra Franz in the tribe Geonemini and distinguished from these genera by the form of the constriction on the head behind the eyes, shape of the nasal plate and median sulcus on the rostrum, and the lack of a tooth on the profemora. Images of the habitus and male genitalia and a diagnosis accompany the description. The new species is named in honor of the late Anne Howden (1927–2016) in recognition of her considerable contribution to our knowledge of Neotropical Entiminae.

The study explores the generic limits and systematic position of Leptoschoinus Dejean, 1836, the type genus for a poorly defined family-group name in Baridinae. Species of five nominal genera, generally with a bipartite denticulation on the endophallus but for historical reasons being assigned to different tribes, were examined for consistency and variants of morphological, mostly male-specific, traits. It is concluded that Leptoschoinus accommodates the most basal elements among the studied species, with plesiomorphic traits dominating. The species of PlatyonyxSchönherr, 1826 and LeptocorynusCasey, 1920 are recognized as advanced members of the same clade, not as a divergent branch of a monophyletic Leptoschoinus. OdontocorynusSchönherr, 1844 is considered as the sister group, with its own similarly inconsistent and progressively diversifying suite of apomorphic states. The following new synonymies are proposed: Leptoschoinus and Leptocorynus are placed in synonymy with Platyonyx; Centrinus nobilis Gistel, 1857 and Leptoschoinus quadriguttatus Voss, 1943 are placed in synonymy with Megabaris quadriguttata (Klug, 1829); Platyonyx ornatus griseofasciatusHustache, 1951 is placed in synonymy with P. ornatusSchönherr, 1826; and Polpones sexmaculatusHustache, 1951 is synonymized with Polpones decemmaculatus Bondar, 1945. Leptocorynus longiclava (Champion, 1908) and Leptocorynus tenuiclava (Champion, 1908) are transferred back to Geraeus Pascoe, 1889 (reinstated combinations). Centrinus curtusHustache, 1951, Leptocorynus aspersus (Champion, 1908), Leptocorynus scutatus (Champion, 1908), and Leptoschoinus fucatus (Klug, 1829) are transferred to Platyonyx (new combinations). Platyonyx ornatus diffususHustache, 1951 is promoted to full species rank (new status). Leptoschoinus quadriguttatus Voss is recognized as a junior secondary homonym of M. quadriguttata (Klug) (new homonymy). The subtribe Leptoschoinina Lacordaire, 1865 is transferred from Madarini Jekel, 1865 to Apostasimerini Schönherr, 1844 (new placement). Platyonyx burkei Prena and Platyonyx wecius Prena are described from central Mexico, and Platyonyx hah Prena is described from Panama, Colombia, and Venezuela (all new species). Previous records of Platyonyx fucatus (Klug) from Mexico apply to P. burkei, records from Panama to P. hah, and records from French Guiana probably to P. ornatus.

Vauricia howdenae Chamorro and Anderson, new genus and new species of Rhinostomina, is described from Malaysia. The new species can be easily distinguished from all other Orthognathini by the uniquely shaped mandibles, the apically wide and medially shallowly emarginate third tarsomere, the subbasally wide pronotum, and the contiguous procoxae, among other characters. We also here retain Yuccaborus LeConte as a distinct genus from Rhinostomus Rafinesque, thus recognizing three genera in the subtribe. An updated key to the world genera of Orthognathini is provided.

Ceutorhynchus howdenae Korotyaev and Anderson, new species (type locality: Canada, North-Western Territories, Fort McPherson) is described. This is the second subarctic species of Ceutorhynchus Germar endemic to North America. It is related to the xerophilous species Ceutorhynchus mutabilis Dietz, to which it is also most similar. The more southern xerophilous C. mutabilis is associated with Erysimum L. (Brassicaceae), and it is possible that C. howdenae uses a host in the same genus, almost certainly in the same family.

Apion carrorum Anderson and Alonso-Zarazaga, new species, is described from Plateau Mountain (50.20741° N, 114.52101° W), Alberta, Canada along the edges of melting snowfields at about 2,400 m elevation. This represents the first description of a species of the genus Apion Herbst in North America. It appears that the species is most similar to Apion arcticum Korotyaev, known only from Wrangel Island in the Russian Far East. Candidate species of Rumex L. as the possible host plant for A. carrorum are discussed.

Macrolechriops moreyi Anzaldo and O'Brien, new species, from Brazil and Uruguay is described and figured. Most examined specimens were reared from the fruits of Maytenus ilicifolia Mart. ex Reissek (Celastraceae), a widely grown shrub with many medicinal uses. The genus MacrolechriopsChampion, 1906 is discussed, and images of type specimens are provided for the other six species currently placed in Macrolechriops to aid future studies on the Lechriopini.

Following up on an inquiry received in 2007 from the late Horace R. Burke, a partial reappraisal of Barber's work on Trichobaris LeConte, 1876 is presented. New information is provided about the composition, identity, and origin of the type material not studied by Barber and, before him, Champion. It is concluded that shortcomings in the access to historical specimens led to misinterpretations and untenable conclusions about species identity. Lectotypes are designated for Baridius trinotatusSay, 1832 and Baridius vestitus Boheman, 1836. The latter name is resurrected from synonymy with Trichobaris trinotata (Say) and takes priority over Trichobaris sororChampion, 1909 (new synonymy). Trichobaris texana LeConte, 1876 is returned into synonymy with Trichobaris pellicea (Boheman, 1844).

Thirteen new species of Laemosaccus Schönherr in two species-groups are described from the eastern and southwestern USA and Baja California: from the USA, Laemosaccus obrieni Hespenheide, new species, Laemosaccus andersoni Hespenheide, new species, Laemosaccus arizonensis Hespenheide, new species, Laemosaccus bimaculatus Hespenheide, new species, Laemosaccus burkei Hespenheide, new species, Laemosaccus clytrinoides Hespenheide, new species, Laemosaccus howdenae Hespenheide, new species, Laemosaccus browerorum Hespenheide, new species, Laemosaccus rileyi Hespenheide, new species, Laemosaccus vaurieae Hespenheide, new species, and Laemosaccus gossypii Hespenheide, new species (nine of these also occur in México); from Baja California, Laemosaccus westcotti Hespenheide, new species and Laemosaccus peninsularis Hespenheide, new species. The previously described Laemosaccus nephele (Herbst) and Laemosaccus texanus Champion are redescribed, and lectotypes are designated for Curculio nephele Herbst and Curculio plagiatus Fabricius. A key is provided for the treated species in genus, and larval and adult host plants are recorded. All but three of the previously undescribed species appear not to have been recognized because of their close resemblance to L. nephele, a putative Batesian mimic of chrysomelids of the subfamily Clytrinae and additional putative Müllerian co-models. That evolutionary selection might produce groups of species difficult to recognize (“mimetic homoplasy”) is discussed and suggested that it may be more frequent in other groups than is currently realized.

Two new species of Isodrusus Sharp are described herein: I. howdenae Cortés-Hernández and Anderson, new species, from Oaxaca, Mexico; and I. curacaoensis Cortés-Hernández and Anderson, new species, from Christoffel National Park, Curaçao, which represents the southernmost occurrence of any described species of the genus. Isodrusus howdenae can be separated from other Isodrusus by the presence of a tooth on the lateral prothoracic margin where the vibrissae are set, the sinuate inner edge of the mesotibiae with two or three small teeth, the lack of a median sulcus on the rostrum, and a small fovea present between the eyes. Isodrusus curacaoensis is distinct in having legs with only one tarsal claw, elytra in lateral view almost evenly arcuate without an abrupt apical declivity, no postocular vibrissae, the antennal scrobe visible in dorsal view, the bowed scape bowed, scales with ecarinate margins, and the anterior margin of the epistome emarginate. A revised key to the species of Isodrusus is provided.

The previously monotypic genus Laccoproctus Schoenherr is revised. In addition to the one known Mexican species, Laccoproctus acalloides Boheman, nine new species are described as follows: Laccoproctus anneae Anderson and O'Brien, new species (Mexico: Guerrero, Hidalgo, Puebla, San Luis Potosí, Veracruz); Laccoproctus burkei Anderson and O'Brien, new species (Mexico: Veracruz, Chiapas); Laccoproctus chiapanecus Anderson and O'Brien, new species (Mexico: Chiapas); Laccoproctus elegantulus Anderson and O'Brien, new species (Mexico: Hidalgo); Laccoproctus nigrinus Anderson and O'Brien, new species (Guatemala); Laccoproctus occidentalis Anderson and O'Brien, new species (Mexico: Colima); Laccoproctus similis Anderson and O'Brien, new species (Mexico: Puebla); Laccoproctus truncatus Anderson and O'Brien, new species (Mexico: Estado de México); and Laccoproctus zunilensis Anderson and O'Brien, new species (Guatemala). A key to the now 10 known species is presented. The genus is proposed as closely related to Aenesias Champion, Lepilius Champion, Enomides Champion, and Plumolepilius Barrios-Izas and Anderson and placed in Molytinae, Conotrachelini (new placement). No detailed information on natural history is available. Specimens of L. elegantulus were collected on trailside ferns at night; specimens of L. chiapanecus were associated with Cedrela odorata L. (‘cedro’); and specimens of other species were collected under bark, on wood, on oak stumps, and on fence posts.

The genus ParanametisBurke, 1960 is revised for North America (including northern Mexico). Paranametis distinctaBurke, 1960 is known from western Texas, New Mexico, and northern Mexico. Phyxelis latirostrisBlatchley, 1916 is resurrected from synonymy with Phyxelis rigidus (Say, 1832) and treated as a valid species transferred to the genus Paranametis as Paranametis latirostris (Blatchley, 1916), resurrected status, new combination. Two new species are described herein: Paranametis burkei Anderson, new species (type locality Big Bend National Park, Brewster County, Texas); and Paranametis howdenae Anderson, new species (type locality Mexico, Nuevo León, 5.3 mi. S La Escondida). All species are described or redescribed, natural history information is summarized, and a listing of locality data from all specimens examined is included and mapped. A key is provided to the species of the genus. All species are represented by habitus images and images of taxonomically significant structures. Paranametis species share a similar ramose/papillate/microfilamentous scale form with species of the genera Phyxelis Schoenherr, 1843 and LeptopinaraO'Brien, 1981, which is unique among North American Byrsopagini.

Phacepholis Horn is a lineage of the Pantomorus-Naupactus complex (Curculionidae: Entiminae: Naupactini) that evolved in North and Central America independently from the species traditionally assigned to Pantomorus Schoenherr. The genus includes 13 species, five distributed in North America and eight in Mexico and Central America. The Mexican-Central American species are Phacepholis globicollis (Pascoe), Phacepholis brevipes (Sharp), Phacepholis trituberculatus (Champion), Phacepholis albicans (Sharp), Phacepholis viridicans (Sharp), Phacepholis strabo (Sharp), Phacepholis howdenae Lanteri and del Río, new species (type locality: Zacapa, Guatemala) and Phacepholis burkei Lanteri and del Río, new species (type locality: Guerrero, Mexico). Phacepholis annectens (Sharp) is a junior synonym of P. brevipes, new synonymy. Lectotypes are designated for Pantomorus brevipes Sharp, Pantomorus annectens Sharp, Pantomorus trituberculatus Champion, Pantomorus albicans Sharp, and Pantomorus viridicans Sharp. Among the eight species of the P. globicollis species-group, six species are endemic to Mexico and two, P. strabo and P. howdenae, range into Central America. The Neotropical Phacepholis mainly differ from the Nearctic species by being larger, with longer antennae, the more convex male pronotum, and the spermathecae subglobose rather than globose. We provide descriptions or redescriptions of the eight species of Phacepholis from Mexico and Central America, a dichotomous key for their identification, habitus photographs, line drawings of male and female genitalia, and distribution maps.

Onthophilus burkei Kovarik and Skelley, new species, is described and illustrated from specimens that were collected in the burrows of Geomys pinetus Rafinesque, the southeastern pocket gopher, in a few scattered localities in Alabama, Georgia, and northern Florida as part of a comprehensive regional effort to sample burrows of this gopher for arthropod inquilines. Onthophilus burkei is the only known burrow-inhabiting Onthophilus from the southeastern USA that has six unmodified pronotal striae. This species appears to be active mainly in winter.

Eulepton burkei Riley, a new genus and new species of eumolpine leaf beetle from the Trans-Pecos Region of western Texas, is described and illustrated. The lspecies' systematic placement is proposed, and a discussion of the unusual character states possessed by the new genus is presented.

Two species of Meloidae from Mexico are described and illustrated. Included are Epicauta (Macrobasis) sorjuanae Pinto, new species (Meloinae: Epicautini) from Oaxaca and Puebla, and Zonitis (Neozonitis) bolognorum Pinto, new species (Nemognathinae: Nemognathini) from Sonora. Placement of each relative to congeners is discussed.

The pubescens species-group of Nearctic Pachybrachis Chevrolat, 1836 is defined and revised. The group is composed of 12 species: P. analis LeConte; P. baja Barney, new species; P. connexus Fall; P. donneri Crotch; P. gilberti Barney, new species; P. morosus Haldeman; P. integratus Fall; P. madera Barney, new species; P. thoracicus Jacoby; P. vigilans Fall; P. wickhami. Bowditch; and P. xanti Crotch. Pachybrachis haematodes Suffrian is synonymized under P. morosus (new synonymy), P. marmoratus Jacoby and P. desertus Fall are synonymized under P. analis (new synonymies), and P. brunneus Bowditch and P. uteanus Fall are synonymized under P. donneri (new synonymies). A key to species is given, and a description, figure, and range map are presented for each species.

The six genera of Agaocephalini in Colombia are distinguished in a dichotomous key and illustrated. Lycomedes enigmaticus Neita-Moreno and Ratcliffe, new species, is described and differentiated by character comparisons with other previously known Colombian species. Lycomedes lydiaeArnaud, 2012 is proposed as a new junior synonym of Lycomedes reichei Brême, 1844.

A new species of the genus Canthidium Erichson from the Los Chimalapas region of Oaxaca, Mexico is described and illustrated. Canthidium kohlmanni Mora-Aguilar and Delgado is diagnosed, and modifications to the key of the North American species of this genus are included. A non-exhaustive list of the endemic fauna and flora from Los Chimalapas region is presented.

Sex identification of many species of adult riffle beetles (Coleoptera: Elmidae) has relied historically upon removal and examination of genitalia, which leads to death if examined individuals are initially alive. This process and its consequences present a substantial problem if investigators intend to study live beetles in captivity, or if an investigator is examining population sex ratios of threatened or at-risk beetles in the wild where destructive sampling should be avoided. We examined external morphological characteristics of three species of Heterelmis Sharp [H. comalensis (Bosse, Tuff, and Brown, 1988), Heterelmis cf. glabra (Horn), and Heterelmis vulnerata (LeConte)] and developed a non-lethal method of identifying sexes by using these characters. Across species, several external morphological characteristics differed between the sexes, but the length of the fifth abdominal ventrite was the most reliable and robust external character found to vary between the sexes. However, sex misidentification error rates using proposed species-specific length thresholds for the fifth ventrite differed among the three Heterelmis species. This easily applied technique has utility for identifying the sexes of the three Heterelmis species in studies that require live subjects and/or minimal mortality, such as in life history studies or observations of sex ratios in the field.

Brodskyella konvickai Ruzzier and Yeshwanth, new species, and Brodskyella viraktamathi Ruzzier and Yeshwanth, new species, from southern India are described. An identification key is provided for the four species of the genus.

Carpelimus koreanus Lee and Ahn, new species, is described from Korea. The new species is significant because it belongs to the transitional group (comprising almost exclusively tropical species) between Thinodromus Kraatz and Carpelimus Leach. A description, habitus photograph, and line drawings of diagnostic characters of the new species are provided to facilitate identification.