Jia Zhang,^1,2,* Lixia Zhao,^3,* Yuting Fu,^1,2,* Fangyuan Liu,^2,4 Zixin Wang,⁵ Yunxia Li,⁵ Gaoping Zhao,^5,6 Wei Sun,^5,6 Baojiang Wu,^1,2 Yongli Song,^1,2 Shaohua Li,³ Chunxia Hao,^1,2 Bilige Wuyun,^1,2 Rihan Wu,^1,2 Moning Liu,⁷ Guifang Cao,⁷ Buhe Nashun,¹ M. Azim Surani,⁸ Qingyuan Sun,⁹ Siqin Bao,^1,2,** Pentao Liu,^10,11,** Xihe Li<sup>1,2,5,**

1</sup>The State Key Laboratory of Reproductive Regulation and Breeding of Grassland Livestock, Inner Mongolia University, Hohhot 010020, China
²Research Center for Animal Genetic Resources of Mongolia Plateau, College of Life Sciences, Inner Mongolia University, Hohhot 010020, China
³College of Basic Medicine, Inner Mongolia Medical University, Hohhot 010110, China
⁴Clinical Medical Research Center, The Affiliated Hospital, Inner Mongolia Medical University, Hohhot 010050, China
⁵Inner Mongolia Saikexing Institute of Breeding and Reproductive Biotechnology in Domestic Animal, Hohhot 011517, China
⁶National Center of Technology Innovation for Dairy Industry, Hohhot 010020, China
⁷Key Laboratory of Basic Veterinary Medicine, College of Veterinary Medicine, Inner Mongolia Agricultural University, Hohhot 010018, China
⁸Wellcome Trust/Cancer Research UK Gurdon Institute, Henry Wellcome Building of Cancer and Developmental Biology, Cambridge CB2 1QN, UK
⁹Fertility Preservation Lab, Reproductive Medicine Center, Guangdong Second Provincial General Hospital, Guangzhou 510317, China
¹⁰School of Biomedical Science, Stem Cell and Regenerative Consortium, Li Ka Shing Faculty of Medicine, The University of Hong Kong, Hong Kong 999077, China
¹¹Centre for Translational Stem Cell Biology, Building 17W, The Hong Kong Science and Technology Park, Hong Kong 999077, China

^*Authors Jia Zhang, Lixia Zhao, and Yuting Fu contributed equally to this work.
^**Correspondence: Siqin Bao, Email: baosq@imu.edu.cn; Pentao Liu, Email: pliu88@hku.hk; Xihe Li, Email: lixh@imu.edu.cn