Yoko Okawara,^1,* Takeshi Sekiguchi,² Aya Ikeda,³ Shingo Miura,³ Hiroshi Sasaki,⁴ Takeshi Fujii,⁵ Yayoi Kaneko<sup>6,**

1</sup>1 Wildlife Conservation Laboratory, Division of Ecosciences, Department of Agriculture, Tokyo Univer
²2 Department of Molecular Biology, Graduate School of Medical Sciences, Kyushu University, 3-1-1 Mai
³3 Laboratory of Wildlife Conservation and Management, School of Human Sciences, Waseda University, M
⁴4 Chikushi Jogakuen University Junior College, 2-12-1 Ishizaka, Dazaifu, Fukuoka 818-0192, Japan
⁵5 Natural Environment Conservation Division, Environment and Citizens Affairs Bureau, Hiroshima Pref
⁶6 Carnivore Ecology and Conservation Research Group, Division of Ecosciences, Department of Agricult

^*Present address: Chemistry, Biology and Marine Sciences Course, Graduate School of Engineering and Science, University of the Ryukyus, Senbaru, Nishiharacho, Okinawa 903-0213, Japan
^**To whom correspondence should be addressed. E-mail: ykaneko7946@gmail.com