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27 March 2013 The Phylogenetic Placement of Oxygaster van Hasselt, 1823 (Teleostei: Cypriniformes: Cyprinidae) and the Taxonomic Status of the Family-Group Name Oxygastrinae Bleeker, 1860
Kevin L. Tang, Daniel N. Lumbantobing, Richard L. Mayden
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Abstract

The cyprinid genus, Oxygaster van Hasselt, 1823, is the name-bearing type genus of the family-group name Oxygastri Bleeker, 1860, which predates many of the commonly used family-group names for the family Cyprinidae. Historically, the phylogenetic position of Oxygaster within the cyprinid phylogeny has been poorly understood, resulting in an uncertain status for its associated family-group name. The phylogenetic placement of Oxygaster and the status of the family-group name Oxygastri are examined herein using sequence data from four loci: two mitochondrial (cytochrome b, cytochrome c oxidase I) and two nuclear (opsin, recombination activating gene 1). A combined data matrix of 4117 bp for 109 taxa was collected and analyzed. Our results recovered Oxygaster within a large and taxonomically disorganized clade, one that has been reported in other studies on the Cyprinidae. The position of Oxygaster has wide-ranging consequences which led us to synonymize a number of cyprinid subfamilies and reorganize the current cyprinid classification. We recognize a monophyletic Oxygastrinae Bleeker, 1860 that includes the following genera: Aphyocypris, Candidia, Ctenopharyngodon, Culter, Hemigrammocypris, Hypophthalmichthys, Ischikauia, Macrochirichthys, Megalobrama, Metzia, Nicholsicypris, Nipponocypris, Ochetobius, Opsariichthys, Oxygaster, Parachela, Paralaubuca, Pararasbora, Parazacco, Squaliobarbus, Xenocyprioides, Xenocypris, Yaoshanicus, and Zacco. We place the following family-group names into the synonymy of Oxygastrinae Bleeker, 1860: Cultrinae Kryzhanovsky, 1947; Hypophthalmichthyinae Günther, 1868; Opsariichthyinae Rendahl, 1928; Squaliobarbinae Howes, 1981; and Xenocypridinae Günther, 1868.

2013 by the American Society of Ichthyologists and Herpetologists
Kevin L. Tang, Daniel N. Lumbantobing, and Richard L. Mayden "The Phylogenetic Placement of Oxygaster van Hasselt, 1823 (Teleostei: Cypriniformes: Cyprinidae) and the Taxonomic Status of the Family-Group Name Oxygastrinae Bleeker, 1860," Copeia 2013(1), 13-22, (27 March 2013). https://doi.org/10.1643/CG-10-121
Received: 3 August 2010; Accepted: 1 September 2012; Published: 27 March 2013
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